PUBLICATION

Regulation of endoderm formation and left-right asymmetry by miR-92 during early zebrafish development

Authors
Li, N., Wei, C., Olena, A.F., and Patton, J.G.
ID
ZDB-PUB-110429-21
Date
2011
Source
Development (Cambridge, England)   138(9): 1817-1826 (Journal)
Registered Authors
Li, Nan, Patton, James G.
Keywords
Kupffer's vesicle, gata5, Left-right asymmetry, miR-92(mir92), zebrafish
MeSH Terms
  • Animals
  • Animals, Genetically Modified
  • Base Sequence
  • Body Patterning/genetics*
  • Cardia/embryology
  • Cardia/metabolism
  • Embryo, Nonmammalian
  • Endoderm/embryology*
  • Endoderm/metabolism
  • GATA5 Transcription Factor/genetics
  • GATA5 Transcription Factor/metabolism
  • Gene Expression Regulation, Developmental
  • MicroRNAs/genetics
  • MicroRNAs/metabolism
  • MicroRNAs/physiology*
  • Tissue Distribution
  • Viscera/embryology
  • Viscera/metabolism
  • Zebrafish/embryology*
  • Zebrafish/genetics
  • Zebrafish/metabolism
PubMed
21447552 Full text @ Development
Abstract
microRNAs (miRNAs) are a family of 21-23 nucleotide endogenous non-coding RNAs that post-transcriptionally regulate gene expression in a sequence-specific manner. Typically, miRNAs downregulate target genes by recognizing and recruiting protein complexes to 3'UTRs, followed by translation repression or mRNA degradation. miR-92 is a well-studied oncogene in mammalian systems. Here, using zebrafish as a model system, we uncovered a novel tissue-inductive role for miR-92 during early vertebrate development. Overexpression resulted in reduced endoderm formation during gastrulation with consequent cardia and viscera bifida. By contrast, depletion of miR-92 increased endoderm formation, which led to abnormal Kupffer's vesicle development and left-right patterning defects. Using target prediction algorithms and reporter constructs, we show that gata5 is a target of miR-92. Alteration of gata5 levels reciprocally mirrored the effects of gain and loss of function of miR-92. Moreover, genetic epistasis experiments showed that miR-92-mediated defects could be substantially suppressed by modulating gata5 levels. We propose that miR-92 is a critical regulator of endoderm formation and left-right asymmetry during early zebrafish development and provide the first evidence for a regulatory function for gata5 in the formation of Kupffer's vesicle and left-right patterning.
Genes / Markers
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Mutations / Transgenics
Human Disease / Model
Sequence Targeting Reagents
Fish
Antibodies
Orthology
Engineered Foreign Genes
Mapping