PUBLICATION

Hand2 ensures an appropriate environment for cardiac fusion by limiting Fibronectin function

Authors
Garavito-Aguilar, Z.V., Riley, H.E., and Yelon, D.
ID
ZDB-PUB-100826-3
Date
2010
Source
Development (Cambridge, England)   137(19): 3215-3220 (Journal)
Registered Authors
Yelon, Deborah
Keywords
Zebrafish, Heart morphogenesis, Lateral plate mesoderm, Hand2, Fibronectin
Datasets
GEO:GSE23381
MeSH Terms
  • Animals
  • Animals, Genetically Modified
  • Basic Helix-Loop-Helix Transcription Factors/deficiency
  • Basic Helix-Loop-Helix Transcription Factors/genetics
  • Basic Helix-Loop-Helix Transcription Factors/metabolism*
  • Cell Polarity
  • Epithelial Cells/cytology
  • Epithelial Cells/metabolism
  • Fibronectins/genetics
  • Fibronectins/metabolism*
  • Gene Expression Regulation, Developmental
  • Heart/embryology*
  • Mutation
  • Myocardium/cytology
  • Myocardium/metabolism*
  • Zebrafish/embryology*
  • Zebrafish/genetics
  • Zebrafish/metabolism*
  • Zebrafish Proteins/deficiency
  • Zebrafish Proteins/genetics
  • Zebrafish Proteins/metabolism*
PubMed
20724450 Full text @ Development
Abstract
Heart formation requires the fusion of bilateral cardiomyocyte populations as they move towards the embryonic midline. The bHLH transcription factor Hand2 is essential for cardiac fusion; however, the effector genes that execute this function of Hand2 are unknown. Here, we provide in zebrafish the first evidence for a downstream component of the Hand2 pathway that mediates cardiac morphogenesis. Although hand2 is expressed in cardiomyocytes, mosaic analysis demonstrates that it plays a non-autonomous role in regulating cardiomyocyte movement. Gene expression profiles reveal heightened expression of fibronectin 1 (fn1) in hand2 mutant embryos. Reciprocally, overexpression of hand2 leads to decreased Fibronectin levels. Furthermore, reduction of fn1 function enables rescue of cardiac fusion in hand2 mutants: bilateral cardiomyocyte populations merge and exhibit improved tissue architecture, albeit without major changes in apicobasal polarity. Together, our data provide a novel example of a tissue creating a favorable environment for its morphogenesis: the Hand2 pathway establishes an appropriate environment for cardiac fusion through negative modulation of Fn1 levels.
Genes / Markers
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Mutations / Transgenics
Human Disease / Model
Sequence Targeting Reagents
Fish
Antibodies
Orthology
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Mapping