PUBLICATION

The Zn Finger protein Iguana impacts Hedgehog signaling by promoting ciliogenesis

Authors
Glazer, A., Wilkinson, A., Backer, C.B., Lapan, S., Gutzman, J.H., Cheeseman, I.M., and Reddien, P.W.
ID
ZDB-PUB-091101-9
Date
2010
Source
Developmental Biology   337(1): 148-156 (Journal)
Registered Authors
Gutzman, Jennifer
Keywords
Planarians, Cilia, Hedgehog, Iguana, Zebrafish
MeSH Terms
  • Adaptor Proteins, Signal Transducing/physiology*
  • Animals
  • Carrier Proteins/physiology*
  • Cilia/physiology*
  • HeLa Cells
  • Hedgehog Proteins/physiology*
  • Humans
  • Planarians/physiology*
  • Signal Transduction/physiology*
  • Zebrafish
  • Zinc Fingers/physiology*
PubMed
19852954 Full text @ Dev. Biol.
Abstract
Hedgehog signaling is critical for metazoan development and requires cilia for pathway activity. The gene iguana was discovered in zebrafish as required for Hedgehog signaling, and encodes a novel Zn finger protein. Planarians are flatworms with robust regenerative capacities and that utilize epidermal cilia for locomotion. RNA interference of Smed-iguana in the planarian S. mediterranea caused cilia loss and failure to regenerate new cilia, but did not cause defects similar to those observed in hedgehog(RNAi) animals. Smed-iguana gene expression was also similar in pattern to the expression of multiple other ciliogenesis genes, but was not required for expression of these ciliogenesis genes. iguana-defective zebrafish had too few motile cilia in pronephric ducts and in Kupffer's vesicle. Kupffer's vesicle promotes left-right asymmetry and iguana mutant embryos had left-right asymmetry defects. Finally, human Iguana proteins (dZIP1 and dZIP1L) localize to the basal bodies of primary cilia and, together, are required for primary cilia formation. Our results indicate that a critical and broadly conserved function for Iguana is in ciliogenesis and that this function has come to be required for Hedgehog signaling in vertebrates.
Genes / Markers
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Mutations / Transgenics
Human Disease / Model
Sequence Targeting Reagents
Fish
Antibodies
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Mapping