PUBLICATION

Proneural gene requirement for hair cell differentiation in the zebrafish lateral line

Authors
Sarrazin, A.F., Villablanca, E.J., Nunez, V.A., Sandoval, P.C., Ghysen, A., and Allende, M.L.
ID
ZDB-PUB-060517-7
Date
2006
Source
Developmental Biology   295(2): 534-545 (Journal)
Registered Authors
Allende, Miguel L.
Keywords
Lateral line, Neuromasts, Hair cells, Proneural genes, bHLH, Atonal homolog, NeuroD
MeSH Terms
  • Animals
  • Basic Helix-Loop-Helix Transcription Factors/genetics*
  • Cell Differentiation/genetics*
  • Cell Lineage
  • Gene Expression Regulation/physiology
  • Hair Cells, Auditory/cytology*
  • Mechanoreceptors*
  • Nerve Tissue Proteins/genetics*
  • Signal Transduction/genetics
  • Transcription Factors/genetics*
  • Zebrafish
  • Zebrafish Proteins/genetics*
PubMed
16678150 Full text @ Dev. Biol.
Abstract
The lateral line system comprises an array of mechanosensory organs, the neuromasts, distributed over the body surface. Each neuromast consists of a patch of mechanosensory hair cells surrounded by support cells. We show that, in the zebrafish, two proneural genes are essential for differentiation of the hair cells, neuroD (nrd) and atonal homolog 1 (ath1). Gene knockdown experiments demonstrate that loss of function of either gene, but not of the related proneural gene neurogenin1 (ngn1), abrogate the appearance of hair cell markers. This is in contrast to other sensory systems, such as the neurons of the lateral line ganglion, where nrd is regulated by ngn1 and not by ath1. Overexpression of ath1 can induce nrd, and the phenotype produced by loss of ath1 function can be partially rescued by injection of nrd mRNA. This supports the conclusion that the activation of nrd probably requires ath1 in the hair cell lineage, whereas in sensory neurons nrd activation requires ngn1. We propose that the emergence of two atonal homologs, ath1 and ngn1, allowed the cellular segregation of mechanoreception and signal transmission that were originally performed by a single cell type as found in insects.
Genes / Markers
Figures
Show all Figures
Expression
Phenotype
Mutations / Transgenics
Human Disease / Model
Sequence Targeting Reagents
Fish
Antibodies
Orthology
Engineered Foreign Genes
Mapping